（二）实际承运人，是指接受承运人委托或者转委托，实际履行本法第四十九条规定的全部或者部分承运人义务的人。
For each minus reaction, the purified DNA fragments were incubated together with three of the four dNTPs, meaning each fragment would be extended by DNA polymerase until the missing nucleotide was needed, at which point synthesis would halt.
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